The imidazole side chain of histidine can function as either a general acid catalyst or a general base catalyst because: A) in the physiological pH range both H+ and OH- are present at high concentrations. B) in the physiological pH range, the nitrogen in the ring can be easily protonated/deprotonated. C) one guanidino group is protonated, and the other is deprotonated at physiological pH. D) the imidazole group is a strong reducing agent at physiological pH. E) the sulfur atoms in the ring can either gain or lose a proton at physiological pH

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